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AJRCMB Masthead

Editor's Office
Editor
Kenneth B. Adler, Ph.D.
North Carolina State University

Deputy Editors
Michael J. Holtzman, M.D.
Washington University School of Medicine

Sadis Matalon, Ph.D.
University of Alabama, Birmingham

ATS Documents Editor
Holger J. Schünemann, M.D., Ph.D.
Italian National Cancer Institute Regina Elena
Rome, Italy

Manager, Peer Review
Karin Beehler

Peer Review Technical Support
Fay Ling

Peer Review Office
(All Submissions)
ATS Peer Review Office
61 Broadway
New York, NY 10006-2755
Tel: (212) 315-8619
Fax: (212) 315-8616

Editorial Production
ATS Editorial Office
61 Broadway, 4th Fl.
New York, NY 10006-2755
Tel: (212) 315-6440
Fax: (212) 315-6456
Internet: http://www.thoracic.org

Publisher
Diane Gern

Editorial Production
Karen Kwak

Business Affairs
Carl S. Aloi, CPA

Subscriptions
Adam Piontek

Advertising Representatives
Cunningham Associates
180 Old Tappan Road
Old Tappan, NJ 07675
Tel: (201) 767-4170
Fax: (201) 767-80

Associate Editors

Zea Borok, M.D.
University
of Southern California

Harold A. Chapman Jr., M.D.
University of California, San Francisco

Augustine M.K. Choi, M.D.
University of Pittsburgh

Gregory P. Downey, M.D.
National Jewish Medical and Research Center

Jack A. Elias, M.D.
Yale University

Patricia Finn, M.D.
University of California, San Diego

Paul S. Foster, Ph.D.
Australian National University, Canberra

Monica Kraft, M.D.
Duke University

Geoffrey Laurent, Ph.D.
University College London, Rayne Institute

George D. Leikauf, Ph.D.
University of Cincinnati

Reynold A. Panettieri, Jr., M.D.
University of Pennsylvania

William C. Parks, Ph.D.
University of Washington School of Medicine

Mark A. Perrella, M.D.
Brigham and Women's Hospital

David A. Schwartz, M.D.
National Jewish Health

Steven D. Shapiro, M.D.
University of Pittsburgh

Hajime Takizawa, M.D., Ph.D.
University of Tokyo Graduate School of Medicine

Judith A. Voynow, M.D.
Duke University

David S. Wilkes, M.D.
Indiana University School of Medicine

Guy A. Zimmerman, M.D.
University of Utah

Editorial Board

Devendra K. Agrawal, Ph.D.                    
Steven M. Albelda, M.D.
Avery August, Ph.D.
Chunxue Bai, M.D.
Zsuzsanna Bebok, M.D.
Tomoko Betsuyaku, M.D.
James C. Bonner, Ph.D.
Arnold R. Brody, Ph.D.
Steven L. Brody, M.D.
G. R. Scott Budinger, M.D.
Edward J. Campbell, M.D.
George H. Caughey, M.D.
David M. Center, M.D.
Rachel C. Chambers, Ph.D.
Edwin Chilvers, Ph.D., F.R.C.P.
Andrew Churg, M.D.
James Collawn, Ph.D.
Erika C. Crouch, M.D., Ph.D.
C. William Davis, Ph.D.
Robert B. Devlin, Ph.D.
Claire M. Doerschuk, M.D.
Jeffrey Drazen, M.D.
Steven M. Dubinett, M.D.
Oliver Eickelberg, M.D.
Serpil Erzurum, M.D.
Bernard M. Fischer, D.V.M., Ph.D.
Rodney J. Folz, M.D.
W. Michael Foster, Ph.D.
Joe G. N. Garcia, M.D.
Jack Gauldie, Ph.D.
Norma Gerard, Ph.D.
Mark N. Gillespie, Ph.D.
Tzipora T. Goldkorn, Ph.D.
Jonathan M. Green, M.D.
Angela Haczku, M.D., Ph.D.
Marc B. Hershenson, M.D.
Stephen T. Holgate, M.D.
John W. Hollingsworth II, M.D.
Patrick G. Holt, D.Sc.
Gary W. Hunninghake, M.D.
David Ingbar, M.D.


Ed Ingenito, M.D., Ph.D.
Charles Irvin, Ph.D.
Yvonne Janssen-Heininger, Ph.D.
Arnold Johnson, Ph.D.
Elliott Kagan, M.D.
Naftali Kaminski, M.D.
Kwang Chul Kim, Ph.D.
Darryl A. Knight, Ph.D.
Lester Kobzik, M.D.
Bev Koller, Ph.D.
Jay K. Kolls, M.D.
Stella Kourembanas, M.D.
Vera Krymskaya, Ph.D.
Wolfgang M. Kuebler, M.D.
Steven L. Kunkel, Ph.D.
Stephen B. Liggett, M.D.
Craig Lilly, M.D.
Andrew H. Limper, M.D.
David A. Lomas, Ph.d., Sc.D.,
 F.R.C.P., F.Med.Sci.
Dwight C. Look, M.D.
Thomas J. Mariani, Ph.D.
William J. Martin II, M.D.
Redwan Moqbel, Ph.D.
Joel Moss, M.D., Ph.D.
Brooke T. Mossman, Ph.D.
Thomas M. Murphy, M.D.
Paul W. Noble, M.D.
Toshihiro Nukiwa, M.D., Ph.D.
Michael A. O'Reilly, Ph.D.
Caroline Owen, M.D., Ph.D.
Linhua Pang, M.D., Ph.D.
Annie Pardo, Ph.D.
R. Stokes Peebles, M.D.
J. Julio Pérez Fontán, M.D.
Bruce Pitt, Ph.D.
Sem H. Phan, M.D.
Marina Pretolani, Ph.D.
Marlene Rabinovitch, M.D.
Usha Raj, M.D.
Scott H. Randall, Ph.D.
Anuradha Ray, Ph.D.



Prabir Ray, Ph.D.
David W.H. Riches, Ph.D.
Karen Ridge, Ph.D.
David M. Rodman, M.D.
Duncan F. Rogers, Sc.D.
Mary C. Rose, Ph.D.
Paul Rothman, M.D.
Matthais Salathe, M.D.
Michael J. Sanderson, Ph.D.
Philip L. Sannes, Ph.D.
Robert Schleimer, Ph.D.
Robert M. Senior, M.D.
Julian Solway, M.D.
Jonathan S. Stamler, M.D.
Theodore J. Standiford, M.D.
Kurt R. Stenmark, M.D.
Robert Strieter, M.D.
Barry R. Stripp, Ph.D.
Jacob I. Sznajder, M.D.
Yohanes Tesfaigzi, M.D.
Victor J. Thannickal, M.D.
Galen B. Toews, M.D.
Rubin Tuder, M.D.
Stefan Uhlig, Ph.D.
Dale T. Umetsu, M.D., Ph.D.
Xiangdong Wang, M.D., Ph.D.
Carl W. White, M.D.
Steve White, M.D.
Jonathan H. Widdicombe, Ph.D.
Marsha A. Wills-Karp, Ph.D.
G. Scott Worthen, M.D.
Joanne Wright, M.D.
Jo Rae Wright, Ph.D.
Reen Wu, Ph.D.
Pan Chyr Yang, M.D., Ph.D.
Joo-Heon Yoon, M.D., Ph.D.
Joseph Zabner, M.D.


The AMERICAN JOURNAL OF RESPIRATORY CELL AND MOLECULAR BIOLOGY (ISSN 1044 - 1549) is published by the American Thoracic Society and issued monthly. The AJRCMB is an official journal of the American Thoracic Society. A volume includes six issues and begins with the January and July issues.

The contents are included in Current Contents/Life Sciences®, Index Medicus®, Medlars®, and Medline®. This journal is the winner of an award of Merit for editorial production and design from the Society for Technical Communication Publications and Art Competition.

Published monthly at 61 Broadway, New York, NY 10006-2755 by the American Thoracic Society. Periodicals postage paid at New York, NY and at additional mailing offices. Postmaster: Send address changes to AJRCMB, 61 Broadway, New York, NY 10006-2755. Printed at Dartmouth Journal Services, Waterbury, VT.

Permission to photocopy for internal or personal use or the internal or personal use of specific clients is granted by the AJRCMB for libraries and other users registered with the Copyright Clearance Center (CCC), provided that the base fee of $2.00 per copy of the artide is paid directly to CCC, 21 Congress St., Salem, MA 01970. Special requests should be addressed to the AJRCMB, Attn: Diane Gern, Publisher, ATS Editorial Office, 61 Broadway, New York, NY 10006-2755.

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